Publications (2014-2016) (An * denotes a non peer reviewed publication or book)


2016 (Including In press papers)

  1. Atkins, S., Cantor, M., Pillay, N., Cliff, G., Keith, M. & Parra,G. (2016). Net loss of endangered humpback dolphins:integrating residency,site and bycatch in sharks nets. Marine Ecology Progress Series. 555:249-260.
  2. Davies, A. B., Marneweck, D.G., Druce, D.J. & Asner, P. (2016). Den site selection, pack composition, and reproductive success in endangered African wild dogs. Behavioural Ecology. 11.doi:10.1093/beheco/arw124.
  3. Dietrich, M., Tjale, M.A., Weyer, J., Kearney, T., Seamark, E.C.J. & Nel, L.H. (2016). Diversity of Bartonella and Rickettsia spp. In Bats and Their Blood-Feeding Ectoparasites from South Africa and Swaziland. Plos One. DOI:10.137/journal.pone.0152077.
  4. Do Linh San, E., Sato, J.J., Belant, J.L. & Somers, M.J. (Eds). In press. Small Carnivores: Evolution, Ecology, Behaviour and Conservation, Wiley-Blackwell, Oxford.
  5. Edwards, S., Aschenborn, O., Gange, A. C., Wiesel, I. (2016). Leopard density estimates from semi-desert commercial farmlands, south-west Namibia. African Journal of Ecology. DOI: 10.1111/aje.12235.
  6. Edwards, S., Gange, A.C., Wiesel, I. (2016). An oasis in the desert: The potential of water sources as camera trap sites in arid environments for surveying a carnivore guild. Journal of Arid Environments 01/2016; 124:304.
  7. Geijzendorffer, I.R., Regan, E. C., Pereira, H. M., Brotons, L., Brummitt, N., Gavish, Y. Walters, M. (2016). Bridging the gap between biodiversity data and policy reporting needs: An Essential Biodiversity Variables perspective. Journal of Applied Ecology. DOI: 10.1111/1365-2664.12417
  8. Hayward, M.W., Ward-Fear, G., Hotellier, F.L. Herman, K., Kabat, A.P. & Gibbons, J.P. (2016). Could biodiversity loss have increased Australia’s bushfire threat? Animal Conservation doi:10.1111/acv.12269.
  9. Isaacs, L., Somers, M.J. & Swanepoel, L. In press. Density of African civets  in a moist mountain bushveld region of South Africa. In: Emmanuel Do Linh San, Jun J. Sato, Jerrold L. Belant & Michael J. Somers (Eds) Small Carnivores: Evolution, Ecology, Behaviour and Conservation, Wiley-Blackwell, Oxford.
  10. Jackson, C.R., Marnewick, K., Lindsey, P.A.,  Roskaft, E. & Robertson, M.P. (2015). Evaluating habitat connectivity methodologies: a case study with endangered African wild dogs in South Africa. Lanscape Ecology. DOI: 10. 1007/s 10980-016-0342-5.
  11. Jolles,  A.E.,  Le Roex, N.,  Flacke, G., Cooper, D., Geoghegan, C. & Somers, M.J. In press. Wildife disease dynamics in carnivore and herbivore hosts in the Hluhluwe-iMfolozi Park. In: Conserving Africa’s mega-diversity in the Anthropocene: the Hluhluwe-iMfolozi Park story. Owen-Smith, N., Archibald, S. & Cromsigt, J.P.G.M. (Eds.). Cambridge University Press, Cambridge.
  12. Karczmarski, L.,  Huang, S. Or, C.K.M.,  Gui, D., Can, S.C.Y.,  Lin, W., Porter, L., Womg, W.,  Zheng, R.,  Ho,Y.,  Chi, S.Y.S.,  Tiongso,  A.J.C.,  Mo, Y.,  Chang, W., Kwok, J.H.W.,  Tang, R.W.K., Lee, A.T.L.,  Yiu, S.,  Keith,  M.  & Gailey, G. Wu. (2016). Humpback Dolphins in Hong Kong and pearl River Delta: Status, Threats and Conservation Challenges. Advances in Marine Biology 73 : 0065-2881.
  13. Kearney, T. C.,  Kieth,  M. & Seamark, E. C. J. In press. New records of bat species using Gatkop Cave in the maternal season. Mammalia doi: 10.1515/mammalia-2015-0043.
  14. Labuschagne, L., Swanepoel, L.H., Taylor, P.J., Belmain, S.R. & Keith, M. Are avian predators effective biological control agents for rodent pest management in agricultural systems? Biological Control 100:94-102.
  15. Lanszki, J.,  Lehoczky, I.,  Kotze,  A. K. & Somers, M.J. (2016). Diet of otters (Lutra lutra) in various habitat types in the Pannonian biogeographical region compared to other regions of Europe. PeerJ 4: e2266.
  16. Miller, J.R.B.,  Balme, G.,  Lindsey, P.A.,  Loveridge, A.J.,  Becker, M.S.,  Begg, C., Brink, H., Dolrenry, S.,  Hunt, J.E.,  Jansson, I.,  MacDonald, D.W.,  Mandisodza-Chikerema, R.L. , Cotterill,A.O.,  Packer, C., Rosengren, D., Straford,  K., Trinkel, M., White, P.A., Winterbach, C., Winterbach, H.E.K. & Funston, P.J. In press. Aging traits and sustainable trophy hunting of African lions. Biological Conservation.
  17. Moore, A.E., Woody Cotterill, F.P.D., Winterbach, C.W., Winterbach, H.E.K.,  Astunes, A. & OˊBrien, S.J. (2016). Genetic Evidence of Contrasting Wetlands and Savannah Habitat Specializations in Different Populations of Lion (Panthera leo). Journal of heredity. doi:10.1093/jhered/esv097
  18. Nortjé, G.P., Van Hoven, V., Laker, M.C., Jordaan, J.C. & Louw, M.A. (2016). Quantifying the impact of off-road driving on root-area distribution in soils. African Journal of Wildlife Research. 46(1):33-48.
  19. Pretorius, Y.,  de Boer, W. F., Kortekaas, K., van Wijngaarden, M., Grant, R. C., Kohi, E. M. &  Prins, H. H. T. (2016). Why elephant have trunks and giraffe long tongues: how plants shape large herbivore mouth morphology? Acta Zoologica.  doi:10.1111/azo.12121
  20. Ramnanan, R., Thorn, M., Tambling, C.J. & Somers, M.J. (In press). Resource partitioning between black-backed jackal and brown hyaena in Waterberg Biosphere Reserve, South Africa. Canid Biology & Conservation. 19(2):8-13.
  21. Ripple, W.J., Chapron, G.  López-bao, J.  Durant, S.M., Macdonald, D.D., Lindsey, P. A., Bennett, E.L.,  Bescha, R.L.,  Bruskotter, J. T. Campos-arceiz, A.  Corlett, R.T.,  Darimont, C.T.,  Dickman, A.J.,  Dirzo, R.,  Dublin, H.T.,  Estes, J.A.,  Everatt, K.T.,  Galetti, M.,  Gosiwam, V.R., Hayward, M.T., Hedges, S., Hoffmann, M., Hunter. L.T.B., Kerley, G.I.H., Letnic, M., Levi, T., Maisels, F., Morrison, J.C.,  Nelson, M.P.,  Newsome, T.M.,  Painter, L.,  Pringle, R.M.,  Sandom, C. J., Terborgh, J., Treves, A., van Valkenburgh, B., Vusetich, J.A. , Wirsing, A.J., Wallach, A.D.,  Wolf, C., Woodroffe, R. ,Young, H.  & Zhang, L. (2016). Saving the worlds Terrestrial Megafauna. BioScience Advance Access.
  22. Rosengren, D., Stratford, K., Trinkel,M.,  White,P,A.,  Winterbach,C.,  Winterbach, H.E.K. & Funston, P.J. (2016). Aging traits and sustainable trophy hunting of African lions. Biological Conservation 201: 160–168.
  23.  Saccaggi, D. L., Karsten, M., Robertson, M. P., Kumschick, S., Somers, M. J., Wilson, J. R. U., & Terblanche, J. S. (2016). Methods and approaches for the management of arthropod border incursions. Biological Invasions 18:1057-1075.
  24. Smith, A. Schoeman, M.C., Keith, M.,  Erasmus, B.F.N., Minadjem, A., Moilanen, A. & Di Minin, E. (2016). Synergistic effects of climate and land-use change on representation of African bats in priority conservation areas. Ecological Indicators 69:276-283.
  25. Somers, M.J.,  Becker, P.,  Druce, D.J.,  Graf, J., Gunther, M., Marneweck, D., Trinkle, M., Moleón, M. & Hayward, M.W. In press. Reassembly of the large predator guild into Hluhluwe-iMfolozi Park (Chapter 12).  Conserving Africa’s mega-diversity in the Anthropocene: the Hluhluwe-iMfolozi Park story. Owen-Smith, N., Archibald, S. & Cromsigt, J.P.G.M. (Eds.). Cambridge University Press, Cambridge.
  26. Tensen, L., Groom, R. J., van Belkom, J., Davies-Mostert, H. T., Marnewick, K., & van Vuuren, B. (2016). Genetic diversity and spatial genetic structure of African wild dogs (Lycaon pictus) in the Greater Limpopo transfrontier conservation area. Conservation Genetics, 17(4): 785–794. article.
  27. van Cakenberghe, V. & Seamark, E. C. J. 2016. Recent changes in African Bat Taxonomy (2015 – 2016). Part I. African Bat Conservation News 41: 3.
  28. van der Waal, C. de Kroon H, van Langevelde F. de Boer,W.F., Heitkonig, I.M.A., Slotow, R. Pretorius,Y. & Prins, H.H.T. (2016). Scale-dependent bi-trophic interactions in a semi-arid savana: how herbivores eliminate benefits of nutrient patchiness to plants. Oecologia 181: 1173–1185.
  29. Van Rooyen, M., van Rooyen, N.,  Nsongola, G., Miabangana, S.E. & Gaugris, J. In press. Floridtic composition, diversity and structure of the forest communities in the Kouilou.
  30. Yui, S.W.,  Parrini, F., Karczmarski, L. & Keith, M. Submitted. In Press. Home range establishment and utilization by reintroduced lions (Panthera leo) in a small South African wildlife reserve. Integrative Zoology.



  1. *ACR. 2015. African Chiroptera Report 2015. AfricanBats, Pretoria. i - xix, 1 - 7001pp. doi: 10.13140/RG.2.1.2201.7762.
  2. *Bothma, J. du P. & du Toit, J.G. 2015. Game Ranch Management, 6th Edition. Van Schaik.
  3. Cersosimo, L. M., Lachance, H., St-Pierre, B., van Hoven, W., & Wright, A.-D. G. (2015). Examination of the Rumen Bacteria and Methanogenic Archaea of Wild Impalas (Aepyceros melampus melampus) from Pongola, South Africa.Microbial Ecology69(3), 577–585. doi:10.1007/s00248-014-0521-3.
  4. Collinson, W. J.,  Parker, D. M.,  Bernard, R. T. F., Reilly, B. K. & Davies-mostert, H. T. (2015). An inventory of vertebrate roadkill in the Greater Mapungubwe Transfrontier Conservation Area , South Africa, African Journal of Wildlife Research 45: 301–311.
  5. Dalerum, F. & Belton, L. (2015). African ungulates recognize a locally extinct native predator. Behavioral Ecology,26(1), 215–222. doi:10.1093/beheco/aru180
  6. Davies-mostert, H. T.,  Mills, M. G. L. & Macdonald, D. W. (2015). The demography and dynamics of an expanding managed African wild dog metapopulation. African Journal of Wildlife Research45: 258–273.
  7. Edwards, S.,  Gange, A. C. & Wiesel, I. (2015). Spatiotemporal resource partitioning of water sources by African carnivores on Namibian commercial farmlands. Journal of Zoology297, 22–31. doi:10.1111/jzo.12248.
  8. Everatt, K. T., Andresen, L. & Somers, M. J. (2015). The influence of prey, pastoralism and poaching on the hierarchical use of habitat by an apex predator. African Journal of Wildlife Research45(2), 187–196.
  9. Hayward M.W., Child M.F.,  Kerley, G.I.,  Lindsey, P.A, Somers, M.J.  &  Burns. B. (2015). Ambiguity in guideline definitions introduces assessor bias and influences consistency in IUCN Red List status assessments. Front. Ecol. Evol. 3:87. doi: 10.3389/fevo.2015.00087
  10. Hayward, M. W., Boitani, L., Burrows, N. D., Funston, P. J., Karanth, K. U., MacKenzie, D. I. & Yarnell, R. W. (2015). Ecologists need robust survey designs, sampling and analytical methods. Journal of Applied Ecology 521: 286-290.
  11. *Hayward, M.W. & Somers. M.J. (2014). Translocations in South Africa: lion reintroductions in perspective. Current Conservation. 8.4 published in 2015 but with 2014 date).
  12. Jacques, H., Reed-Smith, J. & Somers, J. (2015). Aonyx capensis. The IUCN Red List of Threatened Species2015. E.T1793A21938767.
  13. Jordaan, R. K., Mcintyre, T., Somers, M. J. & Bester, M. N. (2015). An assessment of spatial and temporal variation in the diet of Cape clawless otters (Aonyx capensis) in marine environments. African Journal of Wildlife Research 45: 342-353.
  14. *Labuschagne, L. 2015. The Barn Owl Secret weapon in the fight against rodents. Farmers Weekly. 16 October: 26-28.
  15. Leijenaar, S.L., Cilliers, D. & Whitehouse-Tedd, K. 2015  Reduction in Livestock Losses following placement of Livestock Guarding Dogs and the impact of herd species and dog sex.  Journal of Agriculture and Biodiversity Research. 4: 9-15. (ISSN 2277-0836)
  16. Marneweck, D., Cameron, E. Z., Ganswindt, A. & Dalerum, F. (2015). Behavioural and endocrine correlates to the aardwolf mating system. Mammalian Biology - Zeitschrift Für Säugetierkunde, 80(1), 31–38. doi:10.1016/j.mambio.2014.08.001
  17. Marnewick, K. & Somers, M. J. (2015). Home ranges of cheetahs ( Acinonyx jubatus ) outside protected areas in South Africa. African Journal of Wildlife Research45: 223–232.
  18. McManus, J. S.,  Dalton, D. L., Kotzé, A., Smuts, B., Dickman, A., Marshal, J. P. & Keith, M. (2015). Gene flow and population structure of a solitary top carnivore in a human-dominated landscape. Ecology and Evolution5(2), 335–344. doi:10.1002/ece3.1322
  19. Périquet, S., Valeix, M., Claypole, J., Drouet-Hoguet, N., Salnicki, J., Mudimba, S. &  Fritz, H. 2015. Spotted hyaenas switch their foraging strategy as a response to changes in intraguild interactions with lions. Journal of Zoology 297: 245-254.
  20. Pfeiffer, M. B., Venter, J. A. & Downs, C. T. (2015). Foraging range and habitat use by Cape Vulture Gyps coprotheres from the Msikaba colony, Eastern Cape province, South Africa. Koedoe, 57(1): 1–11. doi:10.4102/koedoe.v57i1.1240.
  21. Pfeiffer, M.B., Venter, J.A. & Downs, C.T. (2015). Identifying anthropogenic threats to Cape Vultures Gyps coprotheres using community perceptions in communal farmland, Eastern Cape Province, South Africa. Bird Conservation International, 25: 353-365 doi:10.1017/S0959270914000148
  22. Pitman, R. T., Swanepoel, L. H., Hunter, L., Slotow, R. & Balme, G. A. (2015). The importance of refugia , ecological traps and scale for large carnivore management. Biodiversity & Conservation 24: 1975-1987.
  23. Reed-Smith, Jacques,H. & Somers,M,J. 2015. Hydrictics maculicollis. The IUCN Red List of Threatened Species 2015: e.T12420A936042.
  24. Reisinger, R. R., Keith, M., Andrews, R. D. & de Bruyn, P. J. N. (2015). Movement and diving of killer whales (Orcinus orca) at a Southern Ocean archipelago. Journal of Experimental Marine Biology and Ecology473, 90–102. doi:10.1016/j.jembe.2015.08.008
  25. Somers, M.J. & Webster, A. 2015. On-going mesocarnivore research at Telperion. The Diamond Route Newsletter.June. 6.
  26. Swanepoel, L. H., Somers, M. J. & Dalerum, F. (2015). Density of leopards Panthera pardus on protected and non-protected land in the Waterberg Biosphere, South Africa. Wildlife Biology21(5), 263–268. doi:10.2981/wlb.00108.
  27. Swanepoel, L. H., Somers, M. J., van Hoven, W., Schiess-Meier, M., Owen, C., Snyman, A. &  Dalerum, F. (2015). Survival rates and causes of mortality of leopards Panthera pardus in southern Africa. Oryx49(04), 595–603. doi:10.1017/S0030605313001282
  28. Swanepoel, L.H., Somers, M.J., Dalerum, F. ( 2015). Functional Responses of Retaliatory Killing versus Recreational Sport Hunting of Leopards in South Africa. PLoS ONE 10(4): e0125539. doi:10.1371/journal.pone.0125539.
  29. Thorn, M., Green, M., Marnewick, K. & Scott, D. M. (2015). Determinants of attitudes to carnivores: implications for mitigating human–carnivore conflict on South African farmland. Oryx, 49: 270–277. doi:10.1017/S0030605313000744
  30. Turner, J.A., Vasicek, C.A. & Somers, M.J. 2015. Effects of a colour variant on hunting ability: the white lion in South Africa. Open Science Repository Biology, Online(open-access), p.e45011830. Available at: - See more at:
  31. *VAN CAKENBERGHE, V. & SEAMARK, E. C. J. 2015. Recent changes in African Bat taxonomy (2014-2015). Part I.African Bat Conservation News 38: 4.
  32. *VAN CAKENBERGHE, V. and SEAMARK, E. C. J. 2015b. Recent changes in African Bat taxonomy (2014-2015). Part II.African Bat Conservation News 39: 2.
  33. Vasicek, C. A, & Gaugris, J. Y. (2014). Household Firewood Utilization around the Hlatikhulu Forest Reserve , South Africa. Ethnobotany Research & Applications 12: 597–605. (published in 2015 but with 2014 date)
  34. Weel, S., Watson, L. H., Weel, J. & Venter, J. A. (2015). Cape mountain zebra in the Baviaanskloof Nature Reserve, South Africa : resource use reveals limitations to zebra performance in a dystrophic mountainous ecosystem. Afr. J. Ecol. 53: 428-438.
  35. Weise, F. J., Wiesel, I., Jr, J. L. & Vuuren, R. J. Van. (2015). Evaluation of a conflict-related brown hyaena translocation in central Namibia-implications for the future. African Journal of Wildlife Research 45: 178-186.
  36. Winterbach, C. W., Whitesell, C. & Somers, M. J. (2015). Wildlife Abundance and Diversity as Indicators of Tourism Potential in Northern Botswana. Plos One10(8). doi:10.1371/journal.pone.0135595.
  37. Winterbach, H. E. K., Winterbach, C. W., Boast, L. K., Klein, R. & Somers, M. J. (2015). Relative availability of natural prey versus livestock predicts landscape suitability for cheetahs Acinonyx jubatus in Botswana. PeerJ, doi:10.7717/peerj.1033.
  38. Yiu, S.W., Keith, M., Karczmarski, L. & Parrini, F. (2015). Early post-release movement of reintroduced lions (Panthera leo) in Dinokeng Game Reserve, Gauteng, South Africa. European Journal of Wildlife Research. 61: 861-870.


  1. Andresen, L.,  Everatt, K. & Somers,  M.J. 2014. Use of site occupancy models for targeted monitoring of the cheetah.Journal of Zoology, London. 292(3): 212-220.
  2. Balme, G., Lindsey, P., Swanepoel, L.H. & Hunter, L.T.B. In press. Failure of research to address the rangewide conservation needs of large carnivores: leopards in South Africa as a case study. Conservation Letters, Early online, doi:10.1111/conl.12028
  3. Dalerum,F.& Belton,L. (2014). African ungulates recognize a locally extinct native predator. Behavioral Ecology (2014),00(00),1-8.doi:10.1093/beheco/aru180
  4. Dalerum,F. (2014). Identifying the role of conservation biology for solving the environmental crisis. AMBIO. 43:839-846.
  5. Darnell, A.M., Graf, J.A., Somers, M.J., Slotow, R. &  Szykman Gunther, M. (2014). Space use of African wild dogs in relation to other large carnivores. PloS ONE 9(6): e98846. Download here.
  6. Everatt,  K., Andresen, L. & Somers, M.J. 2014. Trophic scaling and occupancy analysis reveals a lion population  limited by top-down anthropogenic pressure in the  Limpopo National Park, Mozambique. PloS ONE 9(6): e99389.
  7. Gaugris, J. & Vasicek,C,A. (2014). A step closer to local carbon calculations: Growth timescales and linera relationships for sand forest and woodland tree species in Maputalnd, South Africa. Open Journal of Foesrty, 2014,4,223-130.
  8. Gaugris,  J. Vasicek, C. & van Rooyen, M.,W. (2014). Woody vegetation utilisation in tembe Elephant Park,Kwazulu Natal, South Africa. International Journal of Biology. 6(3): 1916-9671
  9. Hayward, M. W., Lyngdoh, S., & Habib, B. (2014). Diet and prey preferences of dholes ( C uon alpinus ): dietary competition within Asia’s apex predator guild. Journal of Zoology294. doi:10.1111/jzo.1217
  10. Hunter, S.C., Gusset, M., Miller, L.J. & Somers, M.J. In press. Space use as an indicator of enclosure appropriateness in African wild dogs (Lycaon pictus). Journal of Applied Animal Welfare Science
  11. Miranda, M. Dalerum, F. & Parrini,F. (2014). Interaction patterns within a multi-herbivore assemblage derived from stable isotopes. Ecological Complexity. 20 (2014) 51-60
  12. Mulero-Pázmány, M. Stolper, R. van essen,L,D. Negro, J.J. & Sassen,T. (2014). Remotely piloted aircraft systems as a Rhinoceros abti-poaching tool in Africa. PLOs ONE 9(1). Doi:10.137/journal.pone.0083873.
  13. Pitman,  R.T.,  Mulvaney, J.,  Paul,  M.,  Ramsay,  P.M., Jooste,  E. & Swanepoel, L.H. In press. Global positioning system-located kills and faecal samples: a comparison of leopard Panthera pardus dietary estimates. Journal of Zoology, London.
  14. Pfeiffer, M.B.,  Venter, J.A.  & Downs, C.T., Identifying anthropogenic threats to cape Vultures Gyp coprotheres using community perceptions in communal farmland, Eastern Cape Province, South Africa. Bird Conservation International. doi:10.1017/S)959270914000148.
  15. Roux, A., Beishuizen, R., Brekelmans, W., Ganswindt, A., Paris, M. & Dalerum, F. In press. Innovative parental care in a myrmecophageous mammal. Acta Ethologica. DOI 10.1007/s10211-013-0157-1.
  16. Sapkota, S. Aryal, A.  Baral,  S.R.,  Hayward , M.W. & Raubenheimer,D. (2014). Economic analysis of electric fencing for mitigation human-wildife conflict in Nepal. Journal of Resources and Ecology. 5(3):237-243.
  17. Swanepoel,  L.H., Lindsey, P., Somers, M.J., van Hoven, W. & Dalerum, F. In press. An evaluation of the sustainability of leopard harvest in South Africa. South African Journal of Wildlife Research.
  18. Swanepoel, L.H., Lindsey, P., Somers, M.J., van Hoven, W. & Dalerum, F. 2014. The relative importance of trophy harvest and retaliatory killing for large carnivores: a case study on South African leopards. South African Journal of Wildlife Research. 44: 115-134.
  19. Swanepoel,  L.,H. Somers, M.J., van Hoven, W. Schiess-Meier, M. Owen,C. Snyman,A. Martins,Q. Senekel,C. Camacho,G. Boshoff,W. & Dalerum,F. (2014) Survival rates and causes of mortality of leopards Panthera pardus in Southern Africa. Fauna and International, Oryx.
  20. Winterbach, H.E.K., Winterbach, C.W. & Somers, M.J. 2014. Landscape suitability in Botswana for the conservation of its six large African carnivores.  PloS ONE 9(6): e100202.


- Author M Somers

Copyright © University of Pretoria 2024. All rights reserved.

FAQ's Email Us Virtual Campus Share Cookie Preferences