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Department of Zoology & Entomology
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Prof. Clarke Scholtz
Clarke Scholtz
E-mail: chscholtz@zoology.up.ac.za
Telephone: +27-12-4203236
Research programme

Scarab Research Group

Scarab Research Group

Departmental students currently under supervision
MSc (research)
  1. Ms. Carmen Jacobs
    The lesser of four evils: comparing the effects of four Avermectins on the reproduction and survival of Euoniticellus intermedius
  2. Ms. Angelika Switala
    Beetle relicts of the Western Cape
  1. Mr. James Harrison
    Systematics of the southern African Leucopholina (Scarabaeidae: Melolonthinae: Melolonthini)
  2. Ms. Karien Labuschagne
    Distribution of Culicoides (Diptera: Ceratoopogonidae) species in South Africa
  3. Ms. Robin Lyle
    A taxonomic revision of the Afrotropical Idiopidae (Araneae: Mygalomorphae).
  4. Mr. Robert Nofemela
    The development of a biological control-based integrated management of Plutella xylostella (L>) (Lepidoptera: Plutellidae)
  1. Dr. Suko Mlambo
    Molecular assessment of African Scarabaeinae
  2. Dr. Power Tshikae
    Local and Regional Ecological Factors Influencing Dung Beetle Assemblage structure across an Environmental gradient in Botswana

Departmental students currently under co-supervision
BSc (Hons)
  1. Mr. Deon Bakkes
    Evaluating the probability of conspecificity hypothesis using Pseudoclimaciella Handschin, 1960 (Neuroptera: Mantispidae)
  1. Mr. Ian Engelbrecht
    Systematics and conservation of trapdoor spiders in Gauteng Province, South Africa
Recent Publications
  1. Davis, A.L.V., van Aarde, R.J., Scholtz, C.H., Guldemond, R.A.R., Fourie, J., Deschodt, C.M. (2013) Is microclimate-driven turnover of dung beetle assemblage structure in regenerating coastal vegetation a precursor to re-establishment of a forest fauna? Journal of Insect Conservation. 17:565-576
  2. Tshikae, B.P., Davis, A.L.V., Scholtz, C.H. (2013) Dung beetle assemblage structure across the aridity and trophic resource gradient of the Botswana Kalahari: patterns and drivers at regional and local scales. Journal of Insect Conservation. 17:623-636
  3. Tshikae, B.P., Davis, A.L.V., Scholtz, C.H. (2013) Species richness - Energy relationships and dung beetle diversity across an aridity and trophic resource gradient. Acta Oecologica. 49:71-82
  4. Dacke, M., Baird, E., Byrne, M., Scholtz, C.H., Warrant, E.J. (2013) Dung Beetles Use the Milky Way for Orientation. Current Biology. 23:1-3
  5. Sole, C.L., Scholtz, C.H. (2013) Phylogeographic pointers to conservation needs: South Africa's flagship dung beetle, Circellium bacchusInsect Conservation and Diversity. DOI: doi: 10.1111/icad.12011
  6. Sole, C.L., Scholtz, C.H., Ball, J.B., Mansell, M.W. (2013) Phylogeny and biogeography of southern African spoon-winged lacewings (Neuroptera: Nemopteridae: Nemopterinae). Molecular Phylogenetics and Evolution. 66:360-368
  1. Davis, A.L.V., van Aarde, R.J., Scholtz, C.H., Guldemond, R.A.R., Fourie, J., Deschodt, C.M. (2012) Is microclimate-driven turnover of dung beetle assemblage structure in regenerating coastal vegetation a precursor to re-establishment of a forest fauna? Journal of Insect Conservation. DOI: 10.1007/s10841-012-9542-8
  2. Snyman, L.P., Ohl, M., Mansell, M.W., Scholtz, C.H. (2012) A revision and key to the genera of Afrotropical Mantispidae (Neuropterida, Neuroptera), with the description of a new genus. Zookeys. 184:67-93
  3. du Toit, C.A., Holter, P., Lutermann, H., Scholtz, C.H. (2012) Role of dung beetle feeding mechanisms in limiting their suitability as hosts for the nematode Spirocerca lupiMedical and Veterinary Entomology. 26:455-457
  4. Davis, A.L.V., Scholtz, C.H., Swemmer, A.M. (2012) Effects of land usage on dung beetle assemblage structure: Kruger National Park versus adjacent farmland in South Africa. Journal of Insect Conservation. 16(3):399-411
  1. Holter, P., Scholtz, C.H. (2011) Re-establishment of Biting Mouthparts in Desert-Living Dung Beetles (Scarabaeidae: Scarabaeinae) Feeding on Plant Litter—Old Structures Reacquired or New Ones Evolved? Journal of Morphology. 272:1007-1016
  2. Mlambo, S., Sole, C.L., Scholtz, C.H. (2011) Phylogeny of the African ball-rolling dung beetle genus Epirinus Reiche (Coleoptera: Scarabaeidae: Scarabaeinae). Invertebrate Systematics. 25:197-207
  3. Strümpher, W.P., Scholtz, C.H. (2011) Description of a new, possibly extinct, species of flightless Trox F. (Coleoptera: Trogidae) from South Africa. Insect Systematics and Evolution. 42(4):337-348
  4. Deschodt, C.M., Davis, A.L.V., Scholtz, C.H. (2011) New dung beetle (Coleoptera: Scarabaeidae: Scarabaeinae) taxa from arid south-western Africa. Insect Systematics and Evolution. 42(3):277-294
  5. Strümpher, W.P., Scholtz, C.H. (2011) New species of Trogidae (Coleoptera: Scarabaeoidea) from Australia. Australian Journal of Entomology. 50:139-143
  6. Sole, C.L., Wirta, H., Forgie, S.A., Scholtz, C.H. (2011) Origin of Madagascan Scarabaeini dung beetles (Coleoptera: Scarabaeidae): dispersal from Africa. Insect Systematics and Evolution. 42:29-40
  1. Brown, J., Scholtz, C.H., Janeau, J.L., Grellier, S. (2010) Dung beetles (Coleoptera: Scarabaeidae) can improve soil hydrological properties. Applied Soil Ecology. 46(1):9-16
  2. Baird, E., Byrne, M.J., Scholtz, C.H., Warrant, E.J., Dacke, M. (2010) Bearing selection in ball-rolling dung beetles: is it constant? Journal of Comparative Physiology A. 196:801-806
  3. Jacobs, C.T., Scholtz, C.H., Escobar, F., Davis, A.L.V. (2010) How might intensification of farming influence dung beetle diversity (Coleoptera: Scarabaeidae) in Maputo Special Reserve (Mozambique)? Journal of Insect Conservation. 14:389-399
  4. Davis, A.L.V., Scholtz, C.H., Kryger, U., Deschodt, C.M., Strümpher, W.P. (2010) Dung beetle assemblage structure in Tswalu Kalahari Reserve: responses to a mosaic of landscape types, vegetation communities, and dung types. Environmental Entomology. 39(3):811-820
  5. Sole, C. .L. ., Scholtz, C. .H. (2010) Did dung beetles arise in Africa? A phylogenetic hypothesis based on five gene regions. Molecular Phylogenetics and Evolution. DOI: 10.1016/j.ympev.2010.04.023
  6. Davis, A.L.V., Scholtz, C.H. (2010) Speciation and evolution of eco-climatic ranges in the intertropical African dung beetle genus, Diastellopalpus van Lansberge. Biological Journal of the Linnean Society. 99:405-423
  1. Holter, P., Scholtz, C.H., Stenseng, L. (2009) Desert detritivory: Nutritional ecology of a dung beetle (Pachysoma glentoni) subsisting on plant litter in arid South African sand dunes. Journal of Arid Environments. 73(12):1090-1094
  2. Scholtz, C.H., Davis, A.L.V., Kryger, U. (2009) Evolutionary biology and conservation of dung beetles. Pensoft Publishers, Sofia and Moscow. :1-544
  3. Strümpher, W.P., Scholtz, C.H. (2009) New species and status changes of small flightless relictual Trox Fabricius from southern Africa (Coleoptera: Trogidae). Insect Systematics and Evolution. 40:71-84
  4. Brettschneider, H., Chimimba, C.T., Scholtz, C.H., Bastos, A.D.S., Bateman, P.W. (2009) The tusked king cricket (Libanasidus vittatus (Kirby, 1899) Anostostomatidae) from South Africa: morphological and molecular evidence suggest two cryptic species. Insect Systematics and Evolution. 40(1):85-103
  5. Deschodt, C.M., Scholtz, C.H., Kryger, U. (2009) Description of the larva and pupa of Ichnestoma stobbiai Holm 1992 (Scarabaeidae: Cetoniinae), a range-restricted species of conservation concern. African Entomology. 17(1):43-50
  1. Kryger, U., Scholtz, C.H. (2008) Phylogeography and Conservation of the Rare South African Fruit Chafer Ichnestoma stobbiai (Coleoptera: Scarabaeidae). Evolutionary Biology from Concept to Application. 11:181-196
  2. Davis, A.L.V., Frolov, A.V., Scholtz, C.H. (2008) The African dung beetle genera. Protea Book House, Pretoria. :1-272
  3. du Toit, C. .A., Scholtz, C. .H., Hyman, W. .B. (2008) Prevalence of the dog nematode Spirocerca lupi in populations of its intermediate dung beetle host in the Tshwane (Pretoria) Metropole, South Africa. Onderstepoort Journal of Veterinary Research. :315-321
  4. Davis, A.L.V., Brink, D.J., Scholtz, C.H., Prinsloo, L.C., Deschodt, C.M. (2008) Functional implications of temperature-correlated colour polymorphism in an iridescent, scarabaeine dung beetle. Ecological Entomology. 33:771-779
  5. Le Roux, E., Scholtz, C.H., Kinahan, A.A., Bateman, P.W. (2008) Pre- and post-copulatory mate selection strategies of an African dung beetle Circellium bacchus (Coleoptera: Scarabaeidae). Journal of Insect Behaviour. 21:111-122
  6. Sole, C.L., Bastos, A.D.S., Scholtz, C.H. (2008) . Intra-specific patterns of mitochondrial variation in natural population fragments of a localised desert dung beetle species, Pachysoma gariepinum (Coleoptera: Scarabaeidae). Journal of Heredity. 99(5):464-475
  7. Davis, A.L.V., Scholtz, C.H., Deschodt, C. (2008) Multi-scale determinants of dung beetle assemblage structure across abiotic gradients of the Kalahari-Nama Karoo ecotone, South Africa. Journal of Biogeography. 35:1465-1480
  8. Tshikae, B.P., Davis, A.L.V., Scholtz, C.H. (2008) Trophic associations of a dung beetle assemblage (Scarabaeidae: Scarabaeinae) in a woodland savanna of Botswana. Environmental Entomology . 37:431-441
  9. Deschodt, C.M., Scholtz, C.H. (2008) Systematics of South African forest-endemic dung beetles: new genera and species of small Canthonini (Scarabaeidae: Scarabaeinae). African Entomology. 16(1):91-106
  1. Scholtz, C.H., Inward, D.J.G., Kerley, M.D. (2007) Description of a new Palaearctic Trox Fabricius species (Coleoptera: Trogidae) from Thailand, with a review of the Trox opacotuberculatus species-group and its biogeography. Insect Systematics and Evolution. 38(1):93-103
  2. Olwoch, J.M., van Jaarsveld, A.S., Scholtz, C.H., Horak, I.G. (2007) Climate change and the genus Rhipicephalus (Acari: Ixodidae) in Africa. Onderstepoort Journal of Veterinary Research. 74:45-72
  3. Brettschneider, H., Chimimba, C.T., Scholtz, C.H., Bateman, P.W. (2007) Review of southern African Anostostomatidae (Orthoptera: Ensifera), with a key to genera. African Entomology. 15(1):45-72
  4. Deschodt, C., Kryger, U., Scholtz, C.H. (2007) New taxa of relictual Canthonini dung beetles (Scarabaeidae: Scarabaeinae) utilizing rock hyrax middens as refuges in south-western Africa. Insect Systematics and Evolution. 38:361-376
  5. Veldtman, R., McGeoch, M.A., Scholtz, C.H. (2007) Fine-scale abundance and distribution of wild silk moth pupae. Bulletin of Entomological Research. 97:15-27
  6. Kryger, U., Sole, C., Tukker, R., Scholtz, C.H. (2007) Biology and ecology of Circellium bacchus (Fabricius 1781) (Coleoptera Scarabaeidae), a South African dung beetle of conservation concern. Tropical Zoology. 19:185-207
  7. Holter, P., Scholtz, C.H. (2007) What do dung beetles eat? Ecological Entomology. 32:690-697
  8. Kryger, U., Deschodt, C., Davis, A.L., Scholtz, C.H. (2007) Effects of cattle treatment with a fluazuron pour-on on survival and reproduction of the dung beetle species Onthophagus gazella (Fabricius). Veterinary Parasitology. 143:380-384
  9. Veldtman, R., McGeoch, M.A., Scholtz, C.H. (2007) Can life-history and defence traits predict the population dynamics and natural enemy responses of insect herbivores? Ecological Enotmology. 32:662-673
  10. Sole, C.L., Bastos, A.D.S., Scholtz, C.H. (2007) Do individual and combined data analyses of molecules and morphology reveal the generic status of Pachysoma MacLeay (Coleoptera: Scarabaeidae)? Insect Systematics and Evolution. 38:311-330
  1. Forgie, S.A., Kryger, U., Bloomer, P., Scholtz, C.H. (2006) Evolutionary relationships among the Scarabaeini (Coleoptera: Scarabaeidae) based on combined molecular and morphological data. Molecular Phylogenetics and Evolution. 40(3):662-678
  2. Kryger, U., Deschodt, C., Davis, A.L., Scholtz, C.H. (2006) Effects of cattle treatment with a cypermethrin/cymiazol spray on survival and reproduction of the dung beetle species Euoniticellus intermedius (Coleoptera: Scarabaeidae). Bulletin of Entomological Research. 96:597-603
  1. Kryger, U., Deschodt, C.M., Scholtz, C.H. (2005) Effects of fluazuron and ivermectin treatment of cattle on the structure of dung beetle communities. Agriculture, Ecosystems and Environment. 105(4):649-656
  2. Davis, A.L.V., Scholtz, C.H., Deschodt, C.M. (2005) A dung beetle survey of selected Gauteng nature reserves: implications for conservation of the provincial scarabaeine fauna. African Entomology. 13(1):1-16
  3. Sole, C.L., Scholtz, C.H., Bastos, A.D.S. (2005) Phylogeography of the Namib Desert dung beetles Scarabaeus (Pachysoma) MacLeay (Coleoptera: Scarabaeidae). Journal of Biogeography. 32:75-84
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Website Links of Importance
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Curriculum Vitae
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